Swarming strategies for cooperative species.

In this paper, we propose a model to investigate the relative efficiency of simple swarming strategies based on the interplay between spontaneous and recruitment-based emigration. We conduct a dynamical study of the model which combines inverse density dependence, saturation effects and induced vs. diffusion-like population transfer. The influence of the most relevant parameters is explored on a systematic basis, and transition values for which qualitative changes occur in the system's behaviour are given. The model is then used to study colonization of a multiple sites environment, as well as confrontation between species featuring different swarming strategies. Simulation results indicate that cooperative organisms should have an interest in evolving recruitment-based emigration. The corresponding population transfer patterns prove more efficient in invading new territories, eliminating competitors in the process. We suggest that this advantage could have promoted a simple form of coordinated swarming in species featuring a primitive type of cooperation.